Leslie Knope is preparing for the "battle royale": a big meeting between all departments where they argue over funding. Ron Swanson's intimidating presence and libertarian beliefs are pivotal for the parks department to succeed in getting more funds over other departments, but he is too busy preparing for his upcoming tax audit served to him by his first ex-wife, Tammy Swanson. Leslie, Andy Dwyer, and April Ludgate help Ron but discover he has few actual receipts, just notes of purchase that he wrote himself and photographs of gentlemen's agreements. He does, however, have large amounts of gold that he has buried all over Pawnee. Ron explains that Tammy I had been present throughout most of his young life: she delivered him as a baby, was his teacher, and she even took his virginity. However, she was cold and controlling, which forever turned Ron off to blonde women after their divorce.
Tammy I is horrified at Ron's recordkeeping and demands access to every part of his life for the audit. Tammy I's strict maternal nature intimidates everyone into following her orders, which impresses April. Over the course of the next week, Tammy I moves back in with Ron and assumes complete control over him. Ron shaves off his moustache, becomes kind, and refuses to do anything without Tammy I's approval. Leslie needs Ron to be his usual gruff self for the battle royale and confronts Tammy I about the audit, which she admits is fake - she decided to take Ron back after learning about all the gold he had.
Leslie asks Ron's second ex-wife Tammy Swanson for help, but even she is scared of Tammy I after she attacked her with acid. They decide to ask Ron's survivalist mother, Tamara - nicknamed Tammy, or Tammy 0 - for help and she agrees. Tammy 0 challenges Tammy I to a "prairie drink-off" of highly alcoholic Swanson family mash liquor, but Tammy 0 says Ron will return to the farm forever if she wins. Leslie enters the drinking contest to save Ron from his mother and ex-wife, but the liquor immediately makes her too drunk to continue. Ron, finally fed up over people controlling him, chugs the jug dry and tells both Tammys to leave him alone.
In a subplot, Tom Haverford asks Ben Wyatt to look over the finances of his company, Entertainment 720. Ben discovers that Tom and Jean-Ralphio Saperstein are wasting large amounts of money on a state-of-the-art office, giving employees high salaries with full benefits, and paying Detlef Schrempf and Roy Hibbert to play basketball all day, despite having no income. They ignore Ben's warnings that the company will go bankrupt in a month, but Tom later apologizes to Ben after discovering that he was right.
In another subplot, Ann Perkins asks Chris Traeger to film a quick Public Service Announcement about diabetes, but the overly-enthusiastic Chris spends all day doing countless takes, making Ann wonder why she even dated him in the first place.
Ben: [on Entertainment 720] I would guess that they would be bankrupt by the end of this sentence.
April: [on Tammy One] She's the cold distant mother I never had. I love her.
Ron: I hope the rest of your day is cool beans.
Leslie: Why do you have so many guns?
Tamara: This is America isn't it?
Tamara: Then I don't have to answer stupid questions while standing on my own property.
Leslie: OK, you know what? Let's cut the crap. Is this audit even real?
Tammy One: In a sense, yes, but in another, truer sense, no, it is not. I want Ronald back. But I had to learn about his finances to make sure my future was protected. I'm impressed. He's acquired quite a bit of gold...
Leslie: You gold digger! You are literally a gold digger!
Leslie: Basically we are being attacked by Godzilla, and to beat Godzilla, we need Mothra. No offense.
Tammy Two: None taken. I'm very flattered.
Tammy Zero: It's time to settle this.
Tammy One: Ah, an old fashioned prairie drink-off.
[Tammy Zero opens a jug of alcohol]
April: Ugh, what's in that jug? It smells like jet fuel!
Ron: That's Swanson family mash liquor. Made from the finest corn ever grown on American soil. It's only legal use is to strip varnish off of speed boats.
Tammy Zero: If you win, he's all yours, and if I win, I bring him back to the farm for good.
Leslie: Wait, what? That wasn't the deal!
Tammy One: Pour it. I'm thirsty.
Leslie: Pour me one too, then. Let me in here! I'm gonna join you and if I win, Ron stays here with us.
Ron: Leslie, no, don't drink that. We use it to burn warts off of the mules!
[Leslie and the Tammys take a shot of the liquor]
Leslie: ...POISON! UGH! I made a mistake. I made a mistake.
[Leslie is drunk on Swanson liquor]
Ron: She's had enough!
Tammy One: That's not how it works. She's out.
April: Wait, I'm subbing in.
Ron: April, don't. That stuff will melt the shell off of a snail.
April: I'm Puerto Rican, I can handle it.
[April takes a swig of the liquor]
April: [spits out liquor in disgust] F**K! OH MY GOD!